Planarian Resources on the Web:   English Version
Part II of ‘A List of Publications on Japanese Turbellarians (2000) --- Including Titles of Publications on Foreign Turbellarians written by the Japanese Authors ---’.
Compiled and Annotated by Masaharu KAWAKATSU, Masayuki TAKAI & Gen-yu SASAKI

Last update: 26-APR-2003

Contact:   (Kawakatsu)    (Takai)    (Sasaki)


General Biology, Teaching Materials and Short Reports
        Elementary, Academic
Encyclopedias, Databases and Resources on the Web
Freshwater Planarians
Land Planarians (Exclusive of New Zealand and Australian species introduced into Europe)
New Zealand and Australian species recently found in Europe
Regeneration and Morphogenesis
International Symposia (Programs, Abstracts, etc.)
Hobby, Notice Boards, etc.
Gen-yu's Files (By Gen-yu Sasaki)
Teaching Materials and Short Notes
Students Guide to Planarian Laboratories
Japanese abstracts of lectures given at Conferences, Meetings, Workshops, etc.
Bipaliids (Popular Scientific Articles)
Book Reviews, Winners, etc.
Hobby, Essays, Games, Songs, etc.


Information on planarians on web has increased steadily since the late 1980's. However, in the series of ‘A List of Publications on Japanese Turbellarians’ (published annually after 1968: 1968-1998 by Kawakatsu; 1999-2000 by Kawakatsu & Takai), URLs of these websites have not been listed as a general rule.

At present, the number of URLs in various research fields on planarians has been incleasing yearly. Beginning with 2001, Sasaki (a new coauthor of this bibliographic series) has been successful in collecting some 170 URLs in the world (82 in Japan and 87 in other countries -- including several English Versions of articles in the Gen-yu's Files).

Web information, abstracts and articles written in Japanese cannot be opened without the use of software for the Japanese language. Moreover, the contents of Japanese websites on planarians are non-academic in most cases except for a few special articles and abstracts of zoological papers.

We have used the following system in filing planarian websites:

1). Planarian Resources on the Web: Japanese Version. By Gen-yu Sasaki.
Opened separately as follows:

This Japanese Version opens with Japanese language software for Japanese-speaking persons. All of the websites (both in Japanese and in foreign languages) are classified into 10 sections. A short Japanese explanation is given for each website.

2). Planarian Resources on the Web: Apendix. (In Japanese.) By Gen-yu Sasaki.
Opened separately as follows:

This Japanese Version also opens with Japanese language software for Japanese-speaking persons. It includes a list of Japanese tales and/or novels mentioning planarians in their titles, texts, figures, etc.

3). Planarian Resources on the Web: English Version. By M. Kawakatsu, M. Takai & Gen-yu Sasaki.
Opened separately as follows:

This English Version is based essentially upon the Japanese Version mentioned above. However, the citation style and arrangement of URLs are quite different from those of the Japanese Version.

i) Some 87 URLs of websites written in Western languages (English, German, French, Portuguese, Italian, Danish, Swedish) are classified into 7 sections and listed. The correct scientific name(s) of triclad planarian species employed in each article is also shown in following "Note".

ii) Some 82 URLs of websites written in the Japanese language are classified into 10 sections and listed. For each article or information listed, a short, liberal translation of the English title is given in brackets; authors' names are also given. These lists may have reference value for Western turbellariologists.

We thank Dr. Taisaku Nogi (Dept. Cytokine Biol., The Forsyth Inst., Boston, Mass., U.S.A) for the collection of some of listed websites. We also thank Dr. Robert W. Mitchell (Bandera, Tx., U.S.A.) for reading a draft of this web article.

We hope to receive additional URLs on turbellarians from all over the world. Your kind assistance and information will be much appreciated as usual.

Contacts: (Sasaki)    (Kawakatsu)


General Biology, Teaching Materials and Short Reports


The Classics of Biology: PLANARIA. By BioMEDIA ASSOCIATES
Note. The species employed is Girardia tigrina.

Planarian. In: The Stream Study -- Macroinvertebrates that are very sensitive to pollution. (Stream Study Website Manager.)

Paulette Planaria (Yucky Worm World). (Discovery Communications Inc.) (
Note. The species employed is Girardia dorotocephala (not doratocephala).

The Planarian Home Page. (Anonym.)

Planaria. (Anonym.)

Planaria. (Kealing Jr. High School, Austin, Tx.)

Our Friend The Planarian. (By B. Conklin, B. Sullivan, and T. Duncan.)

Dugesia sp. (Dept. of Biology, Southwest Texas State Univ., San Marcos, Tx.)
Note. Girardia sp. No native Dugesia species are found in the US.

Free-Living Flatworms. (Pocatello High School Honors Biology Class, Idaho.)

The Freshwater Triclad Flatworms. In: FRESHWATER INVERTEBRATE SURVEY of SUFFOLK (Anonym.)

Platyhelminthes Lesson. In: Mr. Lynch's Science Class Resources.

The Worms. In: Mr. Biology's Biology Web site.

Platyhelminthes. (By Murrieta Valley High School Science - Mr. Jones.)

Platyhelminthes and Echinodermata. In: Adam's Invertebrate Web Page!!!

Platyhelminthes (with moving pictures). (By J. G. Houseman, Ottawa Univ.)
Note. The animal employed seems to be Girardia tigrina.

Turbellaria. In: Introductory Biology Courseware. (By G. Anderson.)

Sample Student Laboratory. Learning and memory in planaria (Dugesia dorotocephala.) An experiment in animal behavior. (Student Sample Lab.)
Note. The species employed is Girardia dorotocephala.


Invertebrate Zoology Labs: Lab Exercise #9 Flatworms I - Class Turbellaria. (By J.-L. Monfraix, Univ. Florida.)

Class Turbellaria (Flatworms). In: Bioassessment of Freshwaters Using Benthic Macroinvertebrates - A Primer, 1999. (By S. M. Mandaville.) Soil & Water Conservation Society of Metro Halifax (SWCSMH) - Homepage.

Class Turbellaria. In: Animal Diversity Web. The Univ. of Michigan Museum of Zoology.

Platyhelminthes. (By B. Tietjen, Bellarmine College, Dept. Biology.)

PHYLUM PLATYHELMINTHES. In: Evolution, Ecology and Biodiversity (VIRTUAL MICROSCOPE). Univ. of Winnipeg Campus Manitoba.
Note. Dugesia sp. employed may be Girardia tigrina.

Lab 4: Acoelomates & Pseudocoelomates. In: Zoo Lab - A Website for Animal Biology, Univ. of Wisconsin.

Phylum Platyhelminthes. In: BIOLOGY ON THE WWW, Univ. of Manitoba.
Note. Dugesia sp. employed is Girardia sp.

Introduction to the Platyhelminthes. In: Museum of Paleontology, U.C.Berkeley.
Note. Dugesia employed may be Girardia tigrina.

Animals. (By E. Goo.)

Kingdom Animalia (Platyhelminthes, Nematoda and Annelida). In Peter Lortz Home Page, North Seattle Community College in Seattle, Washington.
Note. Dugesia mentioned in the text is Girardia.

Les Platthelminthes et les Nématodes. In: Protocoles de labo, Ottawa Univ. [French]
Note. Dugesia mentioned in the text is Girardia.

Meiofauna Photo Galleries. Fluorescence microscopy of turbellarian musculature. (By M. Hooge, Maine Univ.)

Planarians in the Psychology Classroom: Habituation and Instrumental Conditioning Demonstrations. (By C. I. Abramson, D. E. Kirkpatrick, N. Bollinger, R. Odde & S. Lambert, Oklahoma State Univ.)

The Wandtafeln of Rudolph Leuckart (1822-1898). Vermes Platodes -- Turbellaria. Marine Biological Laboratory, Woods Hole, Mass.

Planarian papers published in the 18-19th Centuries in Europe. (By Gen-yu Sasaki.)

Encyclopedias, Databases and Resources on the Web

flatworm. In: ENCYCLOPÆDIA BRITANNICA. Inc.,5716,119831+1+110639,00.html?query=flatworm

flatworm. In: Electric Library's Free Encyclopedia.

Flatworm. In: Microsoft ® Encarta ® Online Encyclopedia 2001

Turbellarian Taxonomic Database. A listing of the taxonomy of turbellarian platyhelminths. (By S. Tyler & L. F. Bush.)

Turbellaria: brief checklist. In: The European Register of Marine Species.

Dugesia japonica and Girardia tigrina. In: Codon Usage Database.

Science: Biology: Zoology: Acoelomates. In: Open Directory Project

Platyhelminthes & Nematoda, together with Acanthocephala, Mesozoa, Nematomorpha, Nemertinea. In: Internet Resource Guide for Zoology. BIOSIS, Publishers of BIOSIS Previews, Zoological Record, and other lifescience database.

Sites en rapport avec la recherche au Laboratoire de Biologie Parasitaire, Protistologie, Helminthologie, MNHN. Sites de Parasitologie. [French]

Turbellaria (67 items). In: A bank of digital resources for teaching biology. BIODIDAC - Turbellaria.

Planaria. In: PubMed. National Center for Biotechnology Information (NCBI).*

Keyword = "planaria". In: M/e/o/s.

Planarian Resources on the Web: English Version. (By M. Kawakatsu, M. Takai & Gen-yu Sasaki.) (The present webpage.)

Webpages on planarians in the Gen-yu's Files: Contents. English Version. (By Gen-yu Sasaki.)

Freshwater Planarians

Plathelmintes, Turbellaria, Tricladida. In: BIOSPELEOLOGIA DEL PIEMONTE. Atlante Fotographico Sistematico. (Anonym.) [Italian]

Dugesia. In: Biology Advising. Dept of Biology, Univ. of Central Oklahoma.
Note. The species employed is Girardia tigrina.

Invertebrate Anatomy. Dugesia. (By R. S. Fox, Lander Univ., Greenwood, Sc.)
Note. The species employed is Girardia tigrina.

Institute of Theoretical and Experimental Biophysics, RAS. Biomuseum - Katalog. Kollektsiya izobrazheniï planariï byla cozdana na baze laboratorii Laboratoriya biofiziki kletki i mezhkletochnykh vzaimodeïstviï ITZB (zav. chl-korr.RAN., professor Chaïlakhyan L. M.) magistrantom PushchGU Il'yasovym Z., studentkoï biofaka MGU Zhukovoï D. i studentom MGAPI (Pushchinskoe otd.) Petrovym A.
Videoregistratsiya provodilas' na ustanovke, vklyuchayushcheï: binokulyarnyï mikroskop MBS-2. videokameru Sony, videograbber FlyVido, i komp'yuter IBM PC Pentium 120. [Russian]
Note. Photos of living specimens of Girardia tigrina (both sexual and asexual races) are found; photos of Dugesia japonica and Dugesia ryukyuensis (by Wataru Yoshida of Hirosaki Univ., Japan) are also shown.

Worms in Space - Mission of Planarians on the Space Shuttle and a Brazilian Rocket Flight. (By V. A. Cassanto (ITA)).

Japanese freshwater planarians in color (Dr. M. Kawakatsu). English Version. (By Gen-yu Sasaki.)

Planarians found in tanks for tropical fishes in Japan. English Version. (By Gen-yu Sasaki.)

Land Planarians (Exclusive of New Zealand and Australian species introduced into Europe)

bipalium.jpg. In: BIOMEDIA MUSEUM, Univ. Paisley, UK.
Note. A color photo of the dorsal view of a living specimen is shown: a blackish animal with a brownish yellow head plate having a blackish central mark; with 7 whitish lateral bands. Externally, the animal is similar to Bipalium rauchi von Graff, 1899, from Singapore (taf. IX, fig. 36; see also von Graff, 1917, taf. XXXIV, fig. 3). The genital anatomy of this species is not known.

Land Planarians. In: House & Landscape Pest Series. (A&M Univ., Dept. Entomology, Texas.)

Image of the week (April 1997, Week 4) Greenhouse Triclad Flatworm Bipalium kewense (Santa Barbara City College, Biological Sciences).


land planarians - Bipalium kewense Moseley and Dolichoplana striata Moseley. Univ. of Florida; Florida Department of Agriculture and Consumer Services.

New Predators and Parasites of Earthrworms in Illinois. In: INHS Reports May-June 2000.

Robert E. Ogren. [A Biography]. (By R. E. Ogren.)
Note. He mentioned for the "Land Planarian Indices Series published by Ogren & Kawakatsu (1987-1992), Ogren, Kawakatsu & Froehlich (1992-1999) and Kawakatsu, Ogren & Froehlich (2000).,.htm

REOgren / Land Planarians / Microscopy / Genealogy. Welcome to my Manila site.


Terrestrial planarians (Platyhelminthes, Tricladida, Terricola) from the Iberian Peninsula: first records of the family Rhynchodemidae, with the description of a new Microplana species. (By E. Metos, G. Giribet & S. Carranza.)
Note. This is a web version of the paper published in Contributions to Zoology, 67 (4): 267-276 (in 2000). Microplana nana Mateos, Giribet et Carranza, 2000, is not listed in the "Land Planarian Indices Series". Cf. Kawakatsu, Ogren & Froehlich, 2000. Bull. Fuji Women's College, (38), II: 89. Our misunderstanding (a false note) will be corrected in the next issue of that series.

European Land Planarians. (By J. Speight under the supervision of Dr. H. D. Jones, Univ. of Manchester, School of Biological Sciences.)

Land planarians (Bipalium) of Sri Lanka. (By Priyantha.)

Bipaliid land planarians recorded in Chinese and Japanese Materia Medica. English Version. (By Gen-yu Sasaki)

A reprint edition of four English abstracts and a single text of land planarian papers published in the SHIBUKITSUBO (1998-2001). (Compiled by G.-Y. Sasaki, H. Murayama & M. Kawakatsu.)

New Zealand and Australian species recently found in Europe

Note. The name of "Artioposthia triangulata" is found in web articles listed below. Since the new genus Arthurdendius Jones et Gerard, 1999, was established, the correct scientific name of that New Zealand worm is Arthurdendius triangulatus (Dendy, 1895). Jones, H. D. & Gerard, B. M., 1999. A new genus and new species of terrestrial planarian (Platyhelminthes; Tricladida; Terricola) from Scotland and emendation of the genus Artioposthia. Jour. Nat. Hist., 33: 387-394. Cf. Ogren, Kawakatsu & Froehlich, 1999 (Bull. Fuji Women's College, (37), II: 94-95).

For "Geoplana sanguinea", the following subspecies classification is now employed.

Australoplana sanguinea (Moseley, 1877)
Australoplana sanguinea sanguinea (Moseley, 1877)
Australoplana sanguinea alba (Dendy, 1891)
Report on the OECD Workshop on Terrestrial Flatworms held at the Zoology Department, Univ. of Canterbury, Christchurch, New Zealand. (By D. V. Alford, D. Boag, P. M. Johns & G. W. Yeates.)
Note. Kawakatsu was a participant of the Workshop. The Report is an Official General Record of the Workshop. For papers presented at that Workshop, see Pedobiologia, 42 (5-6): 385-584 (in 1998).
Cf. Ogren, Kawakatsu & Froehlich, 1998 (Bull. Fuji Women's College, (36), II: 81).

The earthworm-eater planarian, Artioposthia triangulata. In: Guide to information on annelids, and current worm research and researchers [Annelid Home Page] etc. (By G. Read.)

Australasian flatworms in Cornwall. (By C. N. French.)

New Zealand Flat Worm. (By B. Boag, Scotish Crop Research Institute.)

Assessment of the global potential distribution of the predatory land planarian Artioposthia triangulata (Dendy) (Tricladida: Terricola) from ecoclimatic data. Abstract of a paper published in New Zealand Jour. Zool., 22: 311-318 (in 1995). (By B. Boag, K. A. Evans, G. W. Yeates, P. M. Johns & R. Neilson.)

New Zealand Flatworm Arthurdendyus triangulatus. (formerly Artioposthia triangulata). The Central Science Laboratory, Executive Agency of the Ministry of Agriculture Fisheries and Food, UK.

DEN NEWZEALANDSKE FLADORM. (By O. M. Christensen & J. G. Mather, Aarhus Univ., Denmark. [Danish]

Nyazeeländska plattmasken. Växtinspektionen, Jordbruksverket. [Swedish]

Note. The correct indication of the scientific name of Kontikia andersoni is as follows: Kontikia andersoni Jones, 1981. (Parentheses should be removed.)


Planarian Flatworms. In: British Marine Life Study Society Home Page.

Marine Flatworms of the World! Polyclads from Okinawa Water (Japan). (By Wolfgang Seifarth 1997-2001.)
Note. Four genera and 47 species (mostly unidentified spp.) of Polyclads from Okinawa are listed with their color photos.

Regeneration and Morphogenesis

Vade mecum: web site: flatworm; with a movie. An official site of a journal "Developmental Biology", Academic Press, London.

EXEMPLO REGIONAL: Girardia tigrina (Girard, 1850). A regeneração em planárias. Projeto Qualibio - Biologia. In: VIDA: DIVERSIDARDE & UNIDADE, Univ. Federal da Bahia, Brasil. [Portuguese]

INVERTEBRATES. In: "Welcome to the Sánchez Laboratory". The University of Utah, School of Medicine, Dept. of Neurobiology and Anatomy. The earliest known Experimental Analyses of Regeneration in Planarians: the work of Dalyell and Johnson. The Pioneering Planarian Regeneration Experiments of Harriet Randolf.
Note. Four classical names of freshwater planarians are found in the text of this web article. Their correct scientific names are as follows.
Dugesia lugubris (Schmidt, 1861) for "Pl. lugubris";
Girardia tigrina (Girard, 1850) for "Planaria maculata;
Polycelis (Ijimia) felina (Dalyell, 1814) for "P. cornuta" and "P. felina".

Molecular and Genetic Mechanisms of Reparative Regeneration in Planaria. (By S. A. Lyk'm & N. Yu. Usman.) In: Russian Academy of Science, Shemyakin-Occhinnikov's Inst. of Biorganic Chemistry, Laboratory of Structure and Function of Human Genes.

International Symposium "Evolution 2000". Biodiversity in Network of Bioinformation from the Standpoint of Devo-Evo View at Tôkyô, Japan. Approach for the comparative study of gene expression patterns in brain development. (By K. Ikeo, Abstract.)

Searching for the prototypic eye genetic network: Sine oculis is essential for eye regeneration in planarians. (By D. Pineda, J. Gonzalez, P. Callarerts, K. Ikeo, W. J. Gehring & E. Salo, 2000) (PNAS, 97 (9): 4525-4529, in 2000).

Planarian Regeneration In: Laboratory of Genes for Regeneration, Shemyakin & Ovchinikov Institute of Bioorganic Chemistry, Russian Academy of Science. (By Rebrikov, D., Barsova, E., Bulina, M., Bogdanova, E. & Shagin, D.).
Note. Titles of their 8 papers are listed (protocols are available). Girardia tigrina is the correct name for "Dugesia tigrina".

Identitat posicional a planària: Gens i processos cel.lulars implicats en la formació i el manteniment del patró corporal en planàries d'aigua dolça. In: UBWeb, Divisió III, Genètica. [Spanish]
Note. General introduction of Prof. Dr. J. Baguñà's team is given. 'Publicacions' from the Dept. de Genètica are also given (after 1995; 9 sheets).

International Symposia (Programs, Abstracts, etc.)

PROCEEDINGS OF THE 8th ISBT (Brisbane, August 1996). Abstracts of Contributions (in no particular order) to Appear in HYDROBIOLOGIA, End of 1998.
Note. Thirty-six abstracts and 4 titles of lectures given at the Brisbane Symposium. The original papers were printed in Hydrobiologia, vol. 383 (in 1998; Eds. E. Schockaert, N. Watson & J.-L. Justine.)

PROCEEDINGS OF THE IXth ISBT BARCELONA 2000. In: Berg. J. Zool., 137 (Supplement 1) (in 2001; Ed. E. Schockaert.)

FIELDS OF RESEARCH. At the Laboratory of Zoology at the Limburgs Universitair Centrum, Diepenbeek, Belgium.
Note. The abstracts on planarian papers by M. Indeherbergi and C. Vreys are included.

Hobby, Notice Boards, etc.

flat worms. In: Shapes of Nature, Breaking Symmetry in Inorganic and Living Systems.
Note. A reproduction of Escher's Flat worms is found. Kawakatsu has an original book <M. C. Escher, 1959. Grafiek en Tekeningen. Koninklijke Uitzeverij J. J. Tijl NV Zwolle> (a gift material of Dr. C. den Hartog when we had a visit from him in the summer of 1971). 'Platwormen - Flat worms - Plattwürmer - Planarires' is an article No. 62 (192 mm x 161 mm); its explanation is on p. 20. Kawakatsu has also a miniature postcard reproduction of that lithograph (Art Unlimited, Amsterdam) obtained from a souvenior shop of the Netherlands Park in Nagasaki City.
        The figure was also used as a cover photo of the Journal of Biological Psychology, vol. XVI, No. 2 (Dec. 1974; = The Worm Runners' Digest edited by Dr. J. V. McConnell).
        Additionally, the Pier of Mikuni Harbor (ca. 500 m long), Fukui Pref., Honshû, Japan, was designed and constructed in 1885 by Mr. Escher. He also built several buildings of western style.... He is a father of   M. C. Escher (1898-1972). (Cited from the Asahi News Paper, Feb. 13, 1997.)

Planarian. In: Vose Elementary School General Information, Oregon.

Flatworms. In: Sea Slug Forum, Australian Museum.

Some common problems and how to counter them! In: problem solving page, Marine Links UK. (Anonym.)

Planaria Crossing - Comments on Planaria. (Anonym.)
Note. A notice board: Typical American humor.

The Behavioral Effects of Cryopreservation on Dugesia tigrina (Brown Planaria). In: Aaron's Page of Magic, Fun, and Love. (By A. Blackwell).
Note. Can freezing planarians recover??? It contains almost nothing of value.


Gen-yu's Files (By Gen-yu Sasaki)

[Webpages on planarians in the Gen-yu's Files: Contents.] English version is available.

[Cocoons and baby planarians: Dugesia japonica.]

[Very large specimens of Dugesia japonica.]

[Planarians with different faces: Dugesia austroasiatica? and D. japonica.]

[Planarians found in tropical fish tanks at my home.]

[Reproduction in planarians.]

[A brackish water planarian with three blackish patterns: Miroplana trifasciata?]

[A small flatworm with a strange face found in a fish tank: An unidentified species (order Lecithoepitheliata, family Prorhynchidae).]

[Japanese freshwater planarians in color (Dr. M. Kawakatsu).] English version is available.

[Planarians found in tanks for tropical fishes in Japan.] English version is available.

[Planarian papers published in the 18-19th Centuries in Europe.] English version is available.

[Bipaliid land planarians recorded in Chinese and Japanese Materia Medica.] English version is available.

[The basic knowledge of Chinese Materia Medica --- For the understanding of natural hystory in the prescientific age of Japan.]

Planarian resources on the Web. The present English Version is essentially based upon this Japanese Version.

Planarian resources on the Web: Appendix.

Teaching Materials and Short Notes

[A simple experiment and observation of Dugesia japonica] (By I. Hoshino).

[Observations and experiments on freshwater planarians from Ni'igata Pref. for high school students] (By M. Sato).

[Observations on freshwater planarians (Planaria go go!)] (By M. Okano).

[A simple observation of Dugesia japonica] (By M. Akai).

[An observation of Dugesia japonica collected from Mt. Akagi, Gunma Pref.] (By Kunissa).

[A simple regeneration experiment of Dugesia japonica] (Anonym.).

[Planarians collected in a brooklet: children's regeneration observation in the summer vacation] (By Nakasone).

[A study of abnormal eyes in Dugesia japonica] (The Biology Club of Kôchi Prefectural Ôtemae High School, Kôchi Pref., Shikoku).

[Regeneration experiments of Dugesia japonica] (Kure Miyahara High School, Hiroshima Pref.).

[Regeneration experiments of Dugesia japonica] (Kogane High School, Chiba Pref.).

[A simple regeneration experiment of Dugesia japonica] (Mishima Junior High School, Ibaraki City, Ôsaka Pref.).

[A culture and observation of Dugesia japonica] (Biology Educational Association of Tôkyô).

[A simple culture method of Dugesia japonica] (Biology Educational Association of Tôkyô).

[Planarians grown in tanks for freshwater shrimp culture] (Anonym.).

[Dugesia japonica living in clean water biotops] (Kyôto and the Global Environment).

[Three photos of planarians] (Technology and Education, Faculty of Human Science, Yokohama National Univ.).

[General explanation on flatworms] (By Drs. A. and T. K. Hikosaka).

[A counterplan for the removal of diatoms, green and green-blue algae grown in marine water aquaria] (Anonym.).

[Freshwater planarians from the Lake Towada-ko area (by title only)] (Towada Elementary and Junior High School).

[Observations on the movement and regeneration of Dugesia japonica] (Nagaizumi High School, Shizuoka Pref.).

[An observation of the seasonal change of a population size of Dugesia japonica living in a small pond in our school] (Kanzei High School, Okayama Pref.).

[Two reports on the effect of endocrine disturb substance to the fission in Dugesia japonica] (The Science Club of Seta Agricultural and Forestry High School, Gunma Pref.).

[A note on a planarian] (The Biology Club of Yokkaichi High School, Mie Pref.).

[A productiono of VTR for teaching material of high school biology: Planarian regenation, etc.] (By S. Kobayashi; Nagaizumi High School, SHizuoka Pref.).

[Platyhelminthes: A key for the class and orders] (By Nunomura, N., Curator of the Toyama Science Museum, Toyama, Honshû).

[Plathelminthes] (In: Revision of the Red List: The Invertebrates III. An Announcement of the Environmental Agency, Apr. 12, 2000).
Note. This Revised List includes 7 triclad species classified into 2 categories. CR+EN: Ectoplana limuli (Ijima et Kaburaki, 1916); Dugesia izuensis (Katô, 1916); Phagocata papillifera (Ijima et Kaburaki, 1916); Dendrocoelopsis kishidai Kawakatsu, 1978. VU: Phagocata albata Ichikawa et Kawakatsu, 1962; Phagocata suginoi Kawakatsu, 1974; Phagocata tenella Ichikawa et Kawakatsu, 1963. The Revised Version of the Japanese Red Data Book of Endangered Species based upon the Revisid List is now in preparation.

[A color photo of a polyclad species found in Katsu'ura, Bôsô Peninshula, Chiba Pref., Honshû] (By Aquatic Invertebrate Research Institute).

[A list of Polyclads in Japan, with 7 photos in color] (Anonym.; Dr. Noda, H.).

[Stream-dwelling animals found in the T'ailuké (Tailuge) National Park, Hsüan-lan County, Taiwan] (Tséng, Ching-Hsien).[Chinese]
Note. This is a short report written in Chinese language. Occurrence of a "Japan flatworm"(=Dugesia japonica) is mentioned based upon Dr. Lue's (et al.?, 1984) article.

Students Guide to Planarian Laboratories

[A laboratory guide of Dr. Agata's team] (Dept. of Biology, Fac. of Science, Okayama Univ., Okayama Pref.).
[A laboratory guide of Dr. Watanabe's team] (Dept. of Life Science, Fac. of Science, Himeji Institute of Technology, Hyôgo Pref.).

[A Japanese abstract of Dr. Agata's Seminar (June, 2000)] (Biological Inst., Fac. of Science, Okayama Univ., Okayama).

[A laboratory guide of Dr. Hoshi's team] (Lab. of Developmental and Reproductive Biology, Dept. of Chemistry, Keiô-Gijuku Univ., Yokohama).
Note. The hatching of a juvenile from the cocoon of Dugesita ryukyuensis Kawakatsu, 1976, is shown in a photo.

[A Japanese abstract on the gene engineering of planarian nervous system] (By Nakagawa, M., Tazaki, K., Gaudieri, S., Ikeo, K., Gojôbori, T. & Agata, K.).

[A laboratory guide of Dr. Kuroda's team] (Dept. of Molecular and Cellular Neurobiology, Neurobiology Institute of Tokyo, Tôkyô).

[A laboratory guide of the Dept. of Mechanical Systems on Information Technology, Hachinohe Institute of Technology, Hachinohe, Aomori Pref.].

[A laboratory guide of Dr. Ishida's team] (Dept. of Biology, Fac. of Science, Hirosaki Univ., Aomori Pref.).

[Color photos in life of 6 freshwater planarian species collected in the vicinity of Hirosaki] (by Dr. Ishida's team of Hirosaki Univ. Laboratory of Developmental Biology, Department of Biofunctional Science, Faculty of Agriculture and Life Science, Hirosaki Univ., Aomori. Pref.).

[A guide of S. Watanabe's Seminar] (Biopsychological Research Group, Fac. of Literature, Keio-Gijuku Univ., Tôkyô).

Japanese abstracts of lectures given at Conferences, Meetings, Workshops, etc.

[Titles of planarian papers published in the Bulletin of Kyoto Gakugei University, Ser. B, Nos. 3-17 (1953-1961)] (Cited from "Current contents of academic series in Japan"; by titles only).
Note. Titles of 19 papers were incleded: 4 English articles (Okugawa, 1953, 1956, 1957; Ichikawa & Okugawa, 1958); 7 Japanese articles on the fission of "Dugesia gonocephala" (i.e., Dugesia japonica described in 1964) (Okugawa & Kawakatsu, 1954-1958); 8 articles on the vertical distribution of freshwater planarians (Kawakatsu, 1955, 1956, 1958, 1961a, b; Okugawa & Kawakatsu, 1956a, b; Okugawa, Kawkatsu & Tarui, 1955).

[Isolation of stem cells in planarians] (JSPS Research for the Future Program).

[DEAD box family genes in planarians. A lecture given at the First Meeting of the RNA Study Group, Aug. 1-2, 1997] (By Shibata, N., Umesono, Y., Orii, H., Agata, K. & Watanabe, K.).

[A mutation of somatic cells found in regenerated animals from fission pieces in the planarian. A lecture given at the Jap. Soc. of Developmental Biology] (By Watanabe, K.).

[The role of dorso-ventral interaction in the onset of planarian regeneration. A lecture given at the Jap. Soc. of Developmental Biology, 2000] (By Kato, K., Orii, H., Watanabe, K. & Agata, K.).

[Djvlg1 (Dugesia japonica vasa like gene 1) and nanos homologus gene in totipotent somatic stem cells of planarians. A lecture given at the Second Meeting of the RNA Study Group, July 8-10, 1998] (By Shibata, N., Sato, H., Orii, H., Watanabe, K. & Agata, K.).

[Regeneration system in planarians. A lecture given at the Symposium of the Hokkaidô Association of Molecular Biologists, Sept. 19, 2000] (By Agata, K.). "A list of related references" is also available.

[The difference between sexualized worms and sexual worms in the planarian Dugesia ryukyuensis. A lecture given at the 71st Ann. Meet. of the Zool. Soc. of Japan, Tôkyô, Sept. 21-23, 2000] (By Kobayashi, K., Arioka, S. & Hoshi, M.).

[Variety and universality of the genus Dugesia of Japan studied by comparisons of partial 18S rDNA. A lecture given at the 71st Ann. Meet. of the Zool. Soc. of Japan, Tôkyô, Sept. 21-23, 2000] (By Yoshida, W., Kato, C. & Ishida, S.).

[A biopsychological study of the feeding behavior of Dugesia japonica. A title of a lecture given at the 63rd Ann. Meet. of the Psychological Society of Japan, 1999] (By Hishimura, Y.).

Bipaliids (Popular Scientific Articles)

THE KGB FILES [A knowledge in various matters on Japanese bipaliid land planarians: Observations, photos, links, and a bulletin board] (By M. Unita).

[Queer worms those I have not ever seen -- yellowish and blackish specimens of bipaliids] (By K. Hanamura).

[Yellowish and blackish specimens of bipaliid species observed in the vicinity of the Ebisu Station, Tôkyô] (Anonym. = Azabu Gakuen Schools).

[Yellowish bipaliid species] (By H. Imaizumi).

[Bipalium nobile observed at Yanaka, Taitô-ku, Tôkyô] (Anonym.)

[A blackish bipaliid planarian having a pale coloration on the ventral side of the body] (By Y. Kitamura).

[A black bipaliid species from Mt. Hiuchi-gadake, Gunma Pref., Honshû] (Anonym.)

[A land planarian species with a lanceolate head observed in Miyako-jima Island, the Southwest Islands of Japan] (Sea Star = Y. Itô).
Note. A beautiful color photo showing the enlarged body (anterior half) of a specimen from the Miyako locality was received from the collector (Mr. Y. Ito). Judging from the external appearance of the animal (ca. 10cm long and 5mm wide in a creeping state),it may be Platydemus manokwari de Beauchamp, 1962. The occurrence of this exotic species in Miyako-jima Island was already reported by Kawakatsu et al. (1993: Biol. Inland Waters, Nara, pp. 5-14).

[Two bipaliids: Bipalium nobile? from Ichikawa City (Chiba Pref.) and Tôkyô; a black specie from Kamakura City (Kanagawa Pref.)] (Anonym.)

[My lovely spiders; three bipaliid species are also my pets!] (Anonym. = Mirukashi-hime).

[Photographic records of animals living in a garden of my residence in Kawasaki City, near Tôkyô] (Anonym.).

[A short note on planarians found in Toyama City, Toyama Pref.] (By N. nunomura).

[An Essay: A bipaliid planarian with a yellowish coloration and an earthworm] (Anonym.: Blown with the Coriolis' Wind).

[Two questions on land planarians] (Web Question Corner of the Osaka Museum of Natural History, Ôsaka).

[Three questions on land planarians] (Solvers: H. Abe and N. Koizumi).

[A giant land planarian species was found in Ojiya, Nîgata Prefecture, Honshû, Japan] (The Nîgata Nippô, Evening ed., July 24 (Friday), 1998).
Note. This is a selected article in the Web Page of the Nîgata Nippô (July 25). A copy of the original Japanese article with English explanation was distributed by Kawakatsu. Cf. Murayama & Kawakatsu (1999: Shibukitsubo, (20); Kawakatsu & Takai (1999: Bull. Fuji Women's College, (37), II: 80). The species is Bipalium nobile Kawakatsu et Makino, 1982. See also a web article by Sasaki, Takai & Kawakatsu (

[Our appeal - A protection and recovery of the forest ecosystem in the Ogasawara Islands] (By the Forestry and Forest Products Research Institute, Ministry of Agriculture, Forestry and Fisheries, Japan).
Note. A photo of a living specimen of an exotic land planarian species, Platydemus manokwari de Beauchamp, 1962, was given. Cf. Kawakatsu, Okochi, Sato, Ohbayashi, Kitagawa & Totani (1999: Occ. Publ., Biol. Lab. Fuji Women's College, Sapporo, Japan, (32): 1-8).

Book Reviews, Winners, etc.

[A short introduction of Dr. W. J. Gehring's article published in Biohistory, vol. 4 (1) in 1996: "Evolution of Eyes"] (By the JT Biohistory Research Hall).

[Contents of the book published by Teshirogi, W. & Watanabe, K. (eds.): "Morpho-differentiation in Planarians -- From Biological Basis to Gene Manipulation --"] (BIOWEB, Dynacom Co., Ltd.).
Note. Cf. Kawakatsu & Takai (1999: Bull. Fuji Women's College, (37), II: 82-84).

[A book review: Agata, K., 1996. "An Experimental Guide Book for Children: Planarians as an Immortal Animal"] (By S. Ishida).
Note. Cf. Kawakatsu & Takai (1999: Bull. Fuji Women's College, (37), II: 78).

[Two reports on the 1996 Biwako Prize for Ecology. Dr. Oleg A. Timoshkin (Limnological Institute, Russian Academy of Sciences, Irkutsk, Russia) received the Prize] (By Kyoto Shimbun Topics 1997. Jan. 24).
Note. Cf. Kawakatsu & Takai (1999: Bull. Fuji Women's College, (37), II: 83-84.

Hobby, Essays, Games, Songs, etc.

[Kawakatsu's 3 Japanese essays published in the Library News of Fuji Women's College. No. 8 (1979): Profile of Dr. Masaharu Kawakatsu, Prof. of Biology of Fuji Women's College. No. 19 (1984): A note on the family name Kawakatsu. No. 29 (1987): Detective stories and I] (By M. Kawakatsu).

[An Interview with Dr. Kenji Watanabe: Planarians and a View of our Future Study] (Anonym.) In: The WebNews from The Kobe Shimbun (Newspaper) - Himeji Ed. (Jan, 2001?)

[Ms. Yamamoto's Novel entitled 'Planaria' and planarians known as an experimental animal in having a remarkable regeneration power] (Anonym.) In: The WebNews from the Kobe Shimbun (Newspaper)(Feb. 5, 2001).

[Planarians and I: A reminiscent talk of my student age] (By Kobashi).

[My unspotted home town located on the SW foot of Mt. Fuji, Shizuoka Pref. Planarians are found in clean and cold streams] (Takako-chacha).
Note. Cf. ("Very large specimens of Dugesia japonica" in the Gen-yu's Files").

[My friend - planarians] (By NINA's Twinkle Magic).

[My lovely planarian!] (Anonym.).

[ I found planarian in a clean stream (in Tochigi Pref.?)] (Anonym.).

[Is planarian eatable?] (Web Question Corner of the Osaka Museum of Natural History, Ôsaka).

[What is a planarian? A large dummy display of a planaria in a Campus Festival of Nara Women's University (Oct. 24-25, 1998)] (By K. Yoshikawa).

[My illusion about a high regeneration ability of planarians -- an Essay ] (By M. Matsuo).

[Don't drink! I found a PET bottle with living planarians in my home refrigerator] (By M. Yoshinaka).

[Learning in planarians by a cannibalism of learned planarians] (Anonym.).

[An essay: Dr. J. V. McConnell's "memory pill" extracted from learning planarians] (By K. Kimura).

[A song of planarian (RealAudio)] (Anonym.).

[Fling Planarians in the night sky - an original tune mp3 (compressed into lzh: 5MB!)] (By Kommy).

[Planarian game (Flash required)] (Anonym.).

[A Profile of the Theartical Group Planaria] (By the Planaria Profile).

Since: 08-AUG-2001